Distinguished by the following: total elements in dorsal fin 50-58; total elements in anal fin 43-50; pectoral-fin rays 15-20; anal-fin pterygiophores preceding the first haemal spine 3-4; caudal vertebral count 23-24; SL/total length 0.815-0.938; head length/SL 0.161-0.179; predorsal length/SL 0.188-0.218; prepelvic length/SL 0.157-0.175; preanal length/SL 0.308-0.362; four to 16 teeth on outer row of upper jaw; 8-13 teeth on outer row of lower jaw; longitudinal scale rows 69-98.